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Chromodoris cf. alderi, Chromodoris sp. 2, Chromodoris sp. 3, Chromodoris sp. 4, Chromodoris sp. 5, Chromodoris tinctoria,
Chromodoris sp. 6, Chromodoris sp. 7, Chromodoris sp. 8, and Chromodoris sp. 9.
Goniobranchus sp. 04 can be distinguished by the purple tips of white rounded tubercles. It has a white mantle with a red reticulate pattern, and an orange marginal band. The rhinophore clubs are orange, the gills are translucent orange.
This species is Chromodoris sp. 2 in IPN at page 210:
Gosliner, Terrence M., David W. Behrens & Ángel Valdés. 2008. Indo-Pacific Nudibranchs and Sea Slugs. Sea Challengers Natural History Books.
Gig Harbor, Washington. 426pp.
Pretty similar are:
Goniobranchus reticulatus which has commonly no marginal band and differs by distinct tubercles and by its opaque white gill rachis
Goniobranchus sp. 03 which differ by its diffuse reticulate pattern, and lacks the purple tipped tubercles.
It was posted on 22 Feb. 2012 at these websites as Chromodoris sp. 04 Genus: Alder & Hancock, 1855,
I follow the recent article (published: 10 April 2012)
Rebecca Fay Johnson & Terrence M. Gosliner. 2012.
Traditional Taxonomic Groupings Mask Evolutionary History:
A Molecular Phylogeny and New Classification of the Chromodorid Nudibranchs. online at
and move Chromodoris sp. 04 to the reerected ("rescued from synonymy") genus Goniobranchus Pease, 1866, on which Rebecca and Terry state: "This clade includes all of the Indo-Pacific species of Chromodoris that are not part of the black-lined, planar egg mass clade, except Chromodoris alternata and Chromodoris ambiguus."
Bill Rudman's Sea-Slug Forum has all species of this complex as Chromodoris tinctoria.